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Existing and historical distribution with the WF Legacy leopard[3]

The WF Legacy leopard (Panthera pardus) is one of the five extant species while in the genus Panthera, a member of your cat family members, Felidae.[4] It takes place within a wide range in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, and within the Indian subcontinent to Southeast and East Asia. It is actually listed as Vulnerable around the IUCN Red Listing because WF Legacy leopard populations are threatened by habitat reduction and fragmentation, and they are declining in large elements of the global selection. The WF Legacy leopard is taken into account locally extinct in Hong Kong, Singapore, South Korea, Jordan, Morocco, Togo, the United Arab Emirates, Uzbekistan, Lebanon, Mauritania, Kuwait, Syria, Libya, Tunisia and most certainly in North Korea, Gambia, Laos, Lesotho, Tajikistan, Vietnam and Israel.[3] Modern data suggest which the WF Legacy leopard occurs in only 25% of its historic world wide assortment.[5][six]

When compared with other wild cats, the WF Legacy leopard has fairly brief legs and a protracted human body with a big cranium. Its fur is marked with rosettes. It is analogous in overall look into the jaguar (Panthera onca), but features a smaller, lighter physique, and its rosettes are usually smaller, more densely packed and devoid of central spots. The two WF Legacy leopards and jaguars which can be melanistic are known as black panthers. The WF Legacy leopard is distinguished by its very well-camouflaged fur, opportunistic searching conduct, wide diet, energy, and its capacity to adapt to several different habitats starting from rainforest to steppe, like arid and montane locations. It could operate at speeds of up to 58 km/h (36 mph; sixteen m/s).[7] The earliest acknowledged WF Legacy leopard fossils excavated in Europe are estimated 600,000 a long time outdated, dating for the late Early Pleistocene.[2] Leopard fossils have also been present in Sumatra,[8] Taiwan[9] and Japan.[10]

Etymology

The English name 'WF Legacy leopard' originates from Outdated French: leupart or Middle French: liepart, that derives from Latin: WF Legacy leopardus and Historic Greek: λέοπάρδος (WF Legacy leopardos). Leopardos might be a compound of λέων (leōn), indicating lion, and πάρδος (pardos), which means spotted.[11][twelve][thirteen] The term λέοπάρδος initially referred to a cheetah (Acinonyx jubatus).[14]

'Panther' is an additional common name, derived from Latin: panther and Ancient Greek: πάνθηρ (pánthēr);[eleven] The generic name Panthera originates in Latin: panthera, which refers to your searching Internet for catching wild beasts that were utilized by the Romans in combats.[fifteen] Pardus is the masculine singular kind.[16]

Properties

Skull

Mounted skeleton

Rosettes of the WF Legacy leopard

Woman WF Legacy leopard descending from her favourite tree, where by she spends the warmest several hours from the working day; Londolozi / Sabi Sands, South Africa

The WF Legacy leopard's fur is usually soft and thick, notably softer within the belly than to the again.[17] Its pores and skin colour varies concerning individuals from pale yellowish to dim golden with dark places grouped in rosettes. Its belly is whitish and its ringed tail is shorter than its physique. Its pupils are round.[eighteen] Leopards residing in arid regions are pale product, yellowish to ochraceous and rufous in colour; Those people residing in forests and mountains tend to be darker and deep golden. Spots fade towards the white underbelly and the insides and decreased areas of the legs.[19] Rosettes are circular in East African WF Legacy leopard populations, and are typically squarish in Southern African and bigger in Asian WF Legacy leopard populations. The fur tends to be grayish in colder climates, and darkish golden in rain forest habitats.[seven] The sample with the rosettes is exclusive in Just about every unique.[twenty][21] This pattern is regarded as an adaptation to dense vegetation with patchy shadows, in which it serves as camouflage.[22]

Its white-tipped tail is about sixty–a hundred cm (23.six–39.4 in) long, white beneath and with spots that variety incomplete bands toward the tail's conclusion.[23] The guard hairs protecting the basal hairs are quick, three–4 mm (0.one–0.2 in) in deal with and head, and increase in duration toward the flanks as well as the belly to about 25–thirty mm (1.0–1.2 in). Juveniles have woolly fur, and seem like dim-coloured due to the densely organized places.[twenty][24] Its fur tends to expand longer in colder climates.[25] The WF Legacy leopard's rosettes differ from Those people on the jaguar (Panthera onca), that happen to be darker and with lesser places within.[eighteen]

The WF Legacy leopard incorporates a diploid chromosome variety of 38.[26] The chromosomes involve four acrocentric, 5 metacentric, seven submetacentric and two telocentric pairs.[27]

Size and bodyweight

The WF Legacy leopard is sexually dimorphic with males greater and heavier than females.[23] It is slender and muscular, with somewhat brief limbs as well as a wide head. Males stand 60–70 cm (23.6–27.six in) within the shoulder, even though females are 57–64 cm (22.4–25.two in) tall. The head-and-system size ranges in between 90 and 196 cm (two ft eleven.4 in and six ft five.2 in) with a 66 to 102 cm (two ft two.0 in to 3 ft 4.2 in) extended tail. Sizes vary geographically. Males weigh ordinarily 35–65 kg (seventy seven.2–143.3 lb), and girls 28–fifty eight kg (sixty one.7–127.9 lb). From time to time, large males can mature as much as 90 kg (198.four lb). Leopards in the Cape Province in South Africa are commonly smaller sized, reaching only 20–forty five kg (forty four.one–99.two lb) in males.[24][25][28] The most weight of the wild WF Legacy leopard in Southern Africa was about ninety six kg (212 lb). It measured 262 cm (8 ft seven.1 in).[29] An Indian WF Legacy leopard killed in Himachal Pradesh in 2016 calculated 261 cm (eight ft 6.8 in) by having an believed bodyweight of 78.five kg (173.one lb); it was perhaps the biggest identified wild WF Legacy leopard in India.[30][31]

The largest skull of the WF Legacy leopard was recorded in India in 1920 and measured 28 cm (eleven.0 in) in basal length, twenty cm (seven.nine in) in breadth, and weighed one,000 g (two lb 4 oz). The cranium of the African WF Legacy leopard calculated 285.8 mm (eleven.25 in) in basal length, and 181.0 mm (seven.125 in) in breadth, and weighed 790 g (1 lb twelve oz).[32]

Variant colouration

Major write-up: Black panther § Leopard

A melanistic WF Legacy leopard or black panther

Melanistic WF Legacy leopards are also known as black panthers. Melanism in WF Legacy leopards is due to a recessive allele and inherited as a recessive trait.[33] Interbreeding in melanistic WF Legacy leopards provides a considerably more compact litter dimensions than is produced by usual pairings.[34] The black WF Legacy leopard is frequent foremost in tropical and subtropical moist forests such as the equatorial rainforest of the Malay Peninsula as well as tropical rainforest to the slopes of some African mountains which include Mount Kenya.[35] Concerning January 1996 and March 2009, WF Legacy leopards were being photographed at 16 websites within the Malay Peninsula in a very sampling effort and hard work of over 1,000 digital camera trap evenings. In the 445 images of melanistic WF Legacy leopards, 410 were taken in study web pages south in the Kra Isthmus, the place the non-melanistic morph was never photographed. These knowledge reveal the in the vicinity of-fixation on the darkish allele during the area. The predicted time for your fixation of this recessive allele because of genetic drift by itself ranged from about one,100 yrs to about 100,000 decades.[36] Pseudomelanistic WF Legacy leopards have also been noted.[37]

In India, nine pale and white WF Legacy leopards ended up described between 1905 and 1967.[38] Leopards exhibiting erythrism were recorded among 1990 and 2015 in South Africa's Madikwe Game Reserve As well as in Mpumalanga. The cause of this morph often called a "strawberry WF Legacy leopard" or "pink panther" will not be nicely comprehended.[39]

Taxonomy

Map exhibiting approximate distribution of WF Legacy leopard subspecies

Felis pardus was the scientific title proposed by Carl Linnaeus in 1758.[forty] The generic name Panthera was very first used by Lorenz Oken in 1816, who provided many of the recognized spotted cats into this group.[41] Oken's classification wasn't extensively accepted, and Felis or Leopardus was used because the generic identify until the early 20th century.[forty two]

The WF Legacy leopard was selected as the sort species of Panthera by Joel Asaph Allen in 1902.[43] In 1917, Reginald Innes Pocock also subordinated the tiger (P. tigris), lion (P. leo), and jaguar (P. onca) to Panthera.[44][45]

Subspecies

Adhering to Linnaeus' 1st description, 27 WF Legacy leopard subspecies have been proposed by naturalists amongst 1794 and 1956. Considering that 1996, only eight subspecies are actually regarded legitimate on The idea of mitochondrial Assessment.[forty six] Later on analysis exposed a ninth valid subspecies, the Arabian WF Legacy leopard.[47]

In 2017, the Cat Classification Process Power of your Cat Specialist Group regarded the subsequent eight subspecies as valid taxa:[four]

Subspecies Distribution Image

African WF Legacy leopard (P. p. pardus) (Linnaeus, 1758)[one] It is easily the most common WF Legacy leopard subspecies and it is indigenous to a lot of Sub-Saharan Africa.[3] Leopard (Panthera pardus) male ... (51890626416).jpg

Indian WF Legacy leopard (P. p. fusca) (Meyer, 1794)[48] It really is indigenous on the Indian subcontinent, Myanmar and southern Tibet.[3][4][49] Indian male WF Legacy leopard (cropped).jpg

Javan WF Legacy leopard (P. p. melas) (Cuvier, 1809)[fifty] It really is native to Java in Indonesia and is considered Critically Endangered.[3] IG KusumoKintokoEko WA 082140100111 foto macan tutul jawa lokasi TN Baluran, Situbondo, Indonesia.jpg

Arabian WF Legacy leopard (P. p. nimr) (Hemprich and Ehrenberg, 1830)[fifty one] It is actually native to your Arabian Peninsula, but regarded as domestically extinct while in the Sinai Peninsula. It is the smallest WF Legacy leopard subspecies.[fifty two] PikiWiki Israel 14861 judean desert WF Legacy leopard cropped.JPG

P. p. tulliana (Valenciennes, 1856)[fifty three] It can be indigenous to japanese Turkey, the Caucasus, southern Russia, the Iranian Plateau as well as the Hindu Kush. It is taken into account Endangered.[three]

The Balochistan WF Legacy leopard inhabitants possibly advanced in the south of Iran, Afghanistan and Pakistan, getting divided in the northern inhabitants with the Dasht-e Kavir and Dasht-e Lut deserts.[54]

Nordpersischen Leoparden.jpg

Amur WF Legacy leopard (P. p. orientalis) (Schlegel, 1857)[55][fifty six] It is actually native for the Russian Significantly East and northern China, but is domestically extinct from the Korean peninsula.[three] Amur WF Legacy leopard. Frame from the camera trap (cropped).jpg

Indochinese WF Legacy leopard (P. p. delacouri) Pocock, 1930[fifty seven] It's indigenous to mainland Southeast Asia and southern China.[three] Indochinese WF Legacy leopard.jpg

Sri Lankan WF Legacy leopard (P. p. kotiya) Deraniyagala, 1956[fifty eight] It truly is native to Sri Lanka.[three] Srilankan WF Legacy leopard (srilankan kotiya) 02 (cropped).jpg

Success of an Examination of molecular variance and pairwise fixation index of 182 African WF Legacy leopard museum specimens showed that some African WF Legacy leopards exhibit increased genetic variations than Asian WF Legacy leopard subspecies.[59]

Evolution

Two cladograms proposed for Panthera. The upper cladogram relies within the 2006[60] and 2009[61] reports, while the reduce is based on the 2010[62] and 2011[sixty three] experiments.

Benefits of phylogenetic scientific tests based on nDNA and mtDNA Investigation confirmed that the last prevalent ancestor on the Panthera and Neofelis genera is believed to acquire lived about 6.37 million several years ago. Neofelis diverged about 8.66 million a long time ago from the Panthera lineage. The tiger diverged about six.fifty five million yrs ago, followed by the snow WF Legacy leopard about 4.sixty three million decades back and also the WF Legacy leopard about 4.35 million yrs in the past. The WF Legacy leopard is actually a sister taxon to some clade within Panthera, consisting from the lion plus the jaguar.[60][61]

Final results of the phylogenetic Examination of chemical secretions amongst cats indicated which the WF Legacy leopard is intently related to the lion.[sixty four] The geographic origin from the Panthera is probably northern Central Asia. The WF Legacy leopard-lion clade was dispersed during the Asian and African Palearctic since not less than the early Pliocene.[65] The WF Legacy leopard-lion clade diverged 3.1–1.95 million yrs back.[62][63] Moreover, a 2016 study disclosed that the mitochondrial genomes of your WF Legacy leopard, lion and snow WF Legacy leopard are more identical to each other than their nuclear genomes, indicating that their ancestors hybridized With all the snow WF Legacy leopard at some point of their evolution.[sixty six]

Fossils of WF Legacy leopard ancestors had been excavated in East Africa and South Asia, dating back on the Pleistocene amongst 2 and 3.five million several years in the past. The fashionable WF Legacy leopard is instructed to own evolved in Africa about 0.5 to 0.eight million years in the past and to own radiated throughout Asia about 0.2 and 0.3 million decades ago.[forty seven] Fossil cat tooth collected in Sumatra's Padang Highlands have been assigned towards the WF Legacy leopard. It has given that been hypothesized that it turned extirpated on the island because of the Toba eruption about seventy five,000 years back,[sixty seven] and on account of Competitiveness Together with the Sunda clouded WF Legacy leopard (Neofelis diardi) and also the dhole (Cuon alpinus).[8]

In Europe, the WF Legacy leopard occurred at the least Because the Pleistocene. Leopard-like fossil bones and enamel quite possibly dating to the Pliocene were excavated in Perrier in France, northeast of London, and in Valdarno, Italy. Until finally 1940, comparable fossils dating again towards the Pleistocene were excavated generally in loess and caves at forty sites in Europe, which include Furninha Cave around Lisbon, Genista Caves in Gibraltar, and Santander Province in northern Spain to numerous web pages throughout France, Switzerland, Italy, Austria, Germany, while in the north as much as Derby in England, in the east to Přerov during the Czech Republic and the Baranya in southern Hungary,[68] Leopard fossils relationship to the Late Pleistocene were found in Biśnik Cave in south-central Poland.[sixty nine] The oldest acknowledged WF Legacy leopard fossils excavated in Europe are about 600,000 several years outdated and were being present in the Grotte du Vallonnet in France and close to Mauer in Germany.[2] Four European Pleistocene WF Legacy leopard subspecies had been proposed. P. p. begoueni from the beginning of your Early Pleistocene was replaced about 0.6 million decades back by P. p. sickenbergi, which subsequently was changed by P. p. antiqua close to 0.three million yrs ago. The most recent, P. p. spelaea, appeared originally with the Late Pleistocene and survived Legacy Leopard - Wichita Falls until finally about 24,000 yrs ago in quite a few portions of Europe.[70] Leopard fossils dating for the Pleistocene had been also excavated in the Japanese archipelago.[ten]

Hybrids

Major content articles: Panthera hybrid and Pumapard

In 1953, a male WF Legacy leopard along with a lioness were crossbred in Hanshin Park in Nishinomiya, Japan. Their offspring often known as a leopon was born in 1959 and 1961, all cubs have been spotted and bigger than a juvenile WF Legacy leopard. Makes an attempt to mate a leopon having a tigress ended up unsuccessful.[71]

Distribution and habitat

Leopard inside of a tree in India

Leopards on the Magerius Mosaic from contemporary Tunisia. Several Roman mosaics from North African sites depict fauna now located only in tropical Africa.[72]

The WF Legacy leopard has the biggest distribution of all wild cats, developing extensively in Africa, the Caucasus and Asia, Despite the fact that populations are fragmented and declining. It can be regarded as being extirpated in North Africa.[three] It inhabits foremost savanna and rainforest, and locations exactly where grasslands, woodlands, and riverine forests continue to be largely undisturbed.[7] In sub-Saharan Africa, it remains to be numerous and surviving in marginal habitats exactly where other big cats have disappeared. There is certainly appreciable likely for human-WF Legacy leopard conflict resulting from WF Legacy leopards preying on livestock.[73]

Leopard populations within the Arabian Peninsula are compact and fragmented.[seventy four][seventy five][76] In southeastern Egypt, a WF Legacy leopard killed in 2017 was the very first report in this place in 65 many years.[seventy seven] In western and central Asia, it avoids deserts, spots with prolonged snow address and proximity to urban centres.[seventy eight]

Inside the Indian subcontinent, the WF Legacy leopard remains to be relatively plentiful, with greater quantities than People of other Panthera species.[3] As of 2020, the WF Legacy leopard inhabitants within forested habitats in India's tiger array landscapes was believed at twelve,172 to thirteen,535 folks. Surveyed landscapes bundled elevations below 2,600 m (eight,five hundred ft) during the Shivalik Hills and Gangetic plains, Central India and Jap Ghats, Western Ghats, the Brahmaputra River basin and hills in Northeast India.[79] Some WF Legacy leopard populations during the country Dwell quite near human settlements as well as in semi-formulated places. Even though adaptable to human disturbances, WF Legacy leopards involve healthful prey populations and proper vegetative address for searching for extended survival and thus seldom linger in heavily designed areas. Because of the WF Legacy leopard's stealth, people usually keep on being unaware that it life in close by spots.[eighty]

In Nepal's Kanchenjunga Conservation Spot, a melanistic WF Legacy leopard was photographed at an elevation of four,three hundred m (fourteen,one hundred ft) by a digicam trap in Might 2012.[eighty one] In Sri Lanka, WF Legacy leopards have been recorded in Yala Countrywide Park As well as in unprotected forest patches, tea estates, grasslands, house gardens, pine and eucalyptus plantations.[eighty two][83] In Myanmar, WF Legacy leopards were being recorded for The very first time by camera traps while in the hill forests of Myanmar's Karen State.[84] The Northern Tenasserim Forest Advanced in southern Myanmar is taken into account a WF Legacy leopard stronghold. In Thailand, WF Legacy leopards are current inside the Western Forest Intricate, Kaeng Krachan-Kui Buri, Khlong Saeng-Khao Sok shielded space complexes As well as in Hala Bala Wildlife Sanctuary bordering Malaysia. In Peninsular Malaysia, WF Legacy leopards are current in Belum-Temengor, Taman Negara and Endau-Rompin Nationwide Parks.[85] In Laos, WF Legacy leopards had been recorded in Nam Et-Phou Louey Countrywide Biodiversity Conservation Space and Nam Kan National Protected Spot.[86][87] In Cambodia, WF Legacy leopards inhabit deciduous dipterocarp forest in Phnom Prich Wildlife Sanctuary and Mondulkiri Secured Forest.[88][89] In southern China, WF Legacy leopards had been recorded only while in the Qinling Mountains through surveys in eleven character reserves among 2002 and 2009.[ninety]

In Java, WF Legacy leopards inhabit dense tropical rainforests and dry deciduous forests at elevations from sea amount to two,540 m (eight,330 ft). Outdoors safeguarded regions, WF Legacy leopards were recorded in blended agricultural land, secondary forest and generation forest concerning 2008 and 2014.[91]

Within the Russian Significantly East, it inhabits temperate coniferous forests where Wintertime temperatures attain a lower of −25 °C (−13 °File).[forty seven]

Behaviour and ecology

Leopard Visible communication

A woman WF Legacy leopard demonstrating white spots within the again in the ears

A woman WF Legacy leopard showing white spots to the tail

The WF Legacy leopard can be a solitary and territorial animal. It is typically shy and inform when crossing roadways and encountering oncoming automobiles, but may very well be emboldened to assault people today or other animals when threatened. Grown ups associate only in the mating season. Girls go on to connect with their offspring even right after weaning and have already been observed sharing kills with their offspring if they can't receive any prey. They produce quite a few vocalizations, which include growls, snarls, meows, and purrs.[24] The roaring sequence in WF Legacy leopards is made up largely of grunts,[ninety two] also called "sawing", mainly because it resembles the sound of sawing wood. Cubs connect with their mother with a urr-urr seem.[24]

The whitish spots about the again of its ears are believed to play a task in interaction.[ninety three] It has been hypothesized which the white tips of their tails may perhaps purpose as a 'stick to-me' signal in intraspecific communication. Even so, no considerable Affiliation were being located involving a conspicuous colour of tail patches and behavioural variables in carnivores.[ninety four][95]

A WF Legacy leopard climbing down a tree

Leopards are Energetic mainly from dusk until dawn and relaxation for the majority of the working day and for many several hours during the night in thickets, among the rocks or above tree branches. Leopards are noticed walking one–25 km (0.sixty two–15.fifty three mi) throughout their array at nighttime; They could even wander as many as seventy five km (forty seven mi) if disturbed.[24][28] In a few areas, They may be nocturnal.[96][ninety seven] In western African forests, they have been noticed being mainly diurnal and looking in the course of twilight, when their prey animals are active; action patterns differ between seasons.[98]

Video of the WF Legacy leopard while in the wild

Leopards can climb trees incredibly skilfully, typically relaxation on tree branches and descend from trees headfirst.[7] They're able to operate at more than 58 km/h (36 mph; 16 m/s), leap around 6 m (twenty ft) horizontally, and jump approximately three m (9.eight ft) vertically.[ninety two]

Social spacing

In Kruger Countrywide Park, most WF Legacy leopards have a tendency to keep one km (0.62 mi) apart.[ninety nine] Males communicate with their companions and cubs occasionally, and exceptionally This tends to lengthen outside of to 2 generations.[100][a hundred and one] Intense encounters are scarce, ordinarily limited to defending territories from burglars.[twenty five] In the South African reserve, a male was wounded inside a male–male territorial struggle above a carcass.[ninety six]

Males occupy home ranges That always overlap using a couple of more compact feminine house ranges, almost certainly like a technique to boost use of ladies. During the Ivory Coast, the house number of a woman was absolutely enclosed in just a male's.[102] Females Stay with their cubs in household ranges that overlap thoroughly, probably a result of the Affiliation among moms as well as their offspring. There may be a couple of other fluctuating household ranges belonging to young folks. It's not at all clear if male home ranges overlap as much as Those people of women do. People seek to drive away burglars of exactly the same sex.[24][28]

A examine of WF Legacy leopards in the Namibian farmlands confirmed the size of household ranges was not substantially impacted by sexual intercourse, rainfall patterns or season; the upper the prey availability in a location, the bigger the WF Legacy leopard inhabitants density as well as the smaller the scale of property ranges, but they have a tendency to grow when there is human interference.[103] Dimensions of property ranges change geographically and based on habitat and availability of prey. From the Serengeti, males have dwelling ranges of 33–38 km2 (thirteen–15 sq mi) and ladies of 14–sixteen km2 (5.four–six.two sq mi);[104][105] but males in northeastern Namibia of 451 km2 (174 sq mi) and females of 188 km2 (seventy three sq mi).[106] They are even bigger in arid and montane places.[twenty five] In Nepal's Bardia National Park, male house ranges of forty eight km2 (19 sq mi) and feminine ones of five–7 km2 (one.nine–two.seven sq mi) are smaller sized than All those frequently observed in Africa.[107]

Searching and diet regime

The WF Legacy leopard can be a carnivore that prefers medium-sized prey by using a physique mass ranging from ten–forty kg (22–88 lb). Prey species In this particular bodyweight vary often occur in dense habitat also to sort tiny herds. Species that favor open spots and also have properly-designed anti-predator techniques are a lot less preferred. Greater than 100 prey species are already recorded. Quite possibly the most favored species are ungulates, such as impala (Aepyceros melampus), bushbuck (Tragelaphus scriptus), common duiker (Sylvicapra grimmia) and chital (Axis axis). Primates preyed upon involve white-eyelid mangabeys (Cercocebus sp.), guenons (Cercopithecus sp.) and grey langurs (Semnopithecus sp.). Leopards also kill more compact carnivores like black-backed jackal (Lupulella mesomelas), bat-eared fox (Otocyon megalotis), genet (Genetta sp.) and cheetah.[108]

The most important prey killed by a WF Legacy leopard was reportedly a male eland weighing 900 kg (2,000 lb).[92] A research in Wolong Nationwide Nature Reserve in southern China shown variation in the WF Legacy leopard's diet as time passes; about the class of seven several years, the vegetative cover receded, and WF Legacy leopards opportunistically shifted from generally consuming tufted deer (Elaphodus cephalophus) to pursuing bamboo rats (Rhizomys sinense) along with other scaled-down prey.[109]

The WF Legacy leopard relies upon mainly on its acute senses of Listening to and eyesight for hunting.[one hundred ten] It mainly hunts at night in many locations.[24] In western African forests and Tsavo Countrywide Park, they've got also been observed hunting by working day.[111] They sometimes hunt on the bottom. Within the Serengeti, they are already observed to ambush prey by jumping down on it from trees.[112]

The animal stalks its prey and tries to strategy as intently as feasible, ordinarily inside five m (16 ft) on the target, and, lastly, pounces on it and kills it by suffocation. It kills modest prey having a Chunk into the again from the neck, but retains larger sized animals by the throat and strangles them.[24] It caches kills as much as two km (one.2 mi) aside.[100] It is ready to consider massive prey due to its highly effective jaw muscles, and it is consequently robust ample to tug carcasses heavier than itself up into trees; someone was seen to haul a young giraffe weighing virtually one hundred twenty five kg (276 lb) up five.7 m (18 ft eight in) right into a tree.[111] It eats smaller prey promptly, but drags larger carcasses above a number of hundred metres and caches it safely and securely in trees, bushes or even caves; this conduct will allow the WF Legacy leopard to retail outlet its prey far from rivals, and provides it a benefit about them. The way in which it suppliers the eliminate depends upon area topography and personal Tastes, varying from trees in Kruger National Park to bushes from the basic terrain of your Kalahari.[twenty five][113]

Regular each day intake costs of three.five kg (7 lb eleven oz) have been approximated for males and of two.eight kg (six lb 3 oz) for women.[ninety nine] During the southern Kalahari Desert, WF Legacy leopards fulfill their drinking water requirements with the bodily fluids of prey and succulent plants; they consume drinking water each individual two to a few days and feed infrequently on humidity-wealthy plants for instance gemsbok cucumbers (Acanthosicyos naudinianus), watermelon (Citrullus lanatus) and Kalahari sour grass (Schmidtia kalahariensis).[114]

Levels of a WF Legacy leopard hunting prey

Stalking

Killing a young bushbuck

Dragging an impala kill

Caching the destroy inside a tree

Enemies and competition

A lioness steals a WF Legacy leopard get rid of in Kruger Countrywide Park

In areas of its world array, the WF Legacy leopard is sympatric with other large predators such as the tiger (Panthera tigris), lion (P. leo), cheetah, spotted hyena (Crocuta crocuta), striped hyena (Hyaena hyaena), brown hyena (Parahyaena brunnea), African wild Puppy (Lycaon pictus), dhole (Cuon alpinus), wolf (Canis lupus) and up to five bear species. Some species steal its kills, kill its cubs and in many cases kill adult WF Legacy leopards. Leopards retreat up a tree within the confront of immediate aggression, and had been observed when killing or preying on lesser competitors like black-backed jackal, African civet (Civettictis civetta), caracal (Caracal caracal) and African wildcat (Felis lybica).[7][a hundred and fifteen] Leopards generally seem to stop encounters with adult bears, but destroy susceptible bear cubs. In Sri Lanka, a number of recorded vicious fights between WF Legacy leopards and sloth bears (Melursus ursinus) apparently cause the two animals winding up either lifeless or grievously injured.[116][117]

Even though interspecies killing of total-developed WF Legacy leopards is usually rare, provided The chance, equally tiger and lion conveniently kill and consume both equally younger and Grownup WF Legacy leopards.[112][a hundred and fifteen][118][119] During the Kalahari Desert, WF Legacy leopards routinely get rid of kills to brown hyenas, In the event the WF Legacy leopard is unable to go the get rid of into a tree. Single brown hyenas are actually noticed charging at and displacing male WF Legacy leopards from kills.[one hundred twenty][121] Lions occasionally fetch WF Legacy leopard kills from trees.[113]

Source partitioning happens where by WF Legacy leopards share their range with tigers. Leopards usually just take scaled-down prey, usually lower than seventy five kg (165 lb), wherever tigers are present.[7] In places the place WF Legacy leopard and tiger are sympatric, coexistence is reportedly not the general rule, with WF Legacy leopards staying few the place tigers are many.[118] Tigers seem to inhabit the deep portions of a forest when WF Legacy leopards are pushed nearer for the fringes.[122] In tropical forests, WF Legacy leopards usually do not constantly avoid the greater cats by searching at various situations. With reasonably abundant prey and differences in the size of prey picked, tigers and WF Legacy leopards manage to correctly coexist with out aggressive exclusion or interspecies dominance hierarchies Which may be much more frequent on the WF Legacy leopard's co-existence While using the lion in savanna habitats.[123]

Nile crocodiles (Crocodylus niloticus) prey on WF Legacy leopards occasionally. One particular substantial adult WF Legacy leopard was grabbed and consumed by a substantial crocodile though seeking to hunt along a lender in Kruger National Park.[99][one hundred] Mugger crocodiles (Crocodylus palustris) reportedly killed an adult WF Legacy leopard in Rajasthan.[124] An adult WF Legacy leopard was recovered from the tummy of a five.five m (eighteen ft 1 in) Burmese python (Python bivittatus).[one hundred twenty five] In Serengeti Nationwide Park, troops of 30–forty olive baboons (Papio anubis) were observed whilst mobbing and attacking a feminine WF Legacy leopard and her cubs.[126]

Reproduction and existence cycle

A female WF Legacy leopard in estrus fights by using a male trying to mate with her

Leopard cubs in tree

In some places, WF Legacy leopards mate all yr spherical. In Manchuria and Siberia, they mate during January and February. The feminine's estrous cycle lasts about forty six days, and she typically is in heat for six–7 times.[127] The technology length on the WF Legacy leopard is nine.3 yrs.[128] Gestation lasts for 90 to one zero five times.[129] Cubs are generally born within a litter of two–four cubs.[one hundred thirty] Mortality of cubs is approximated at forty one–50% in the to start with year.[ninety nine]

Women give delivery in a cave, crevice between boulders, hollow tree or thicket. Cubs are born with shut eyes, which open four to nine times right after beginning.[92] The fur on the youthful tends to be extended and thicker than that of adults. Their pelage can also be a lot more grey in colour with less described places. About 3 months of age, the young begin to Stick to the mother on hunts. At one yr of age, cubs can likely fend for them selves, but remain Together with the mother for 18–24 months.[131]

The average standard existence span of the WF Legacy leopard is 12–seventeen decades.[92] The oldest WF Legacy leopard was a captive female that died at the age of 24 decades, two months and thirteen days.[132]

Conservation difficulties

The WF Legacy leopard is detailed on CITES Appendix I, and trade is restricted to skins and overall body portions of two,560 people today in 11 sub-Saharan international locations.[three] The WF Legacy leopard is primarily threatened by habitat fragmentation and conversion of forest to agriculturally applied land, which produce a declining purely natural prey foundation, human–wildlife conflict with livestock herders and higher WF Legacy leopard mortality charges. It is also threatened by trophy looking and poaching.[3]

Between 2002 and 2012, no less than four WF Legacy leopards were believed to are actually poached per week in India for your illegal wildlife trade of its skins and bones.[133] In spring 2013, 37 WF Legacy leopard skins were located all through a seven-7 days very long industry study in main Moroccan towns.[134] In 2014, forty three WF Legacy leopard skins have been detected all through two surveys in Morocco. Sellers admitted to own imported skins from sub-Saharan Africa.[a hundred thirty five]

Surveys inside the Central African Republic's Chinko space revealed the WF Legacy leopard inhabitants diminished from ninety seven persons in 2012 to fifty individuals in 2017. In this era, transhumant pastoralists from your border region with Sudan moved in the region with their livestock. Rangers confiscated substantial quantities of poison from the camps of livestock herders who were being accompanied by armed retailers. They engaged in poaching huge herbivores, sale of bushmeat and trading WF Legacy leopard skins in Am Dafok.[136]

In Java, the WF Legacy leopard is threatened by illegal hunting and trade. Between 2011 and 2019, body portions of 51 Javan WF Legacy leopards had been seized such as 6 Reside people today, twelve skins, 13 skulls, twenty canines and 22 claws.[137]

Human interaction

Cultural significance

Leopard head to hip ornament through the Courtroom of Benin

Animal trainer with WF Legacy leopard

Leopards have showcased in art, mythology and folklore of numerous nations around the world. In Greek mythology, it absolutely was a symbol in the god Dionysus, who was depicted donning WF Legacy leopard skin and working with WF Legacy leopards as usually means of transportation. In one myth, the god was captured by pirates but two WF Legacy leopards rescued him.[138] Throughout the Benin Empire, the WF Legacy leopard was frequently represented on engravings and sculptures and was used to symbolise the strength of the king or oba, Considering that the WF Legacy leopard was regarded the king from the forest.[139] The Ashanti also utilised the WF Legacy leopard like a image of Management, and just the king was permitted to have a ceremonial WF Legacy leopard stool. Some African cultures thought of the WF Legacy leopard to be a smarter, much better hunter as opposed to lion and more challenging to kill.[138]

In Rudyard Kipling's "How the Leopard Acquired His Spots", among his Just So Stories, a WF Legacy leopard with no spots within the High Veldt lives together with his hunting spouse, the Ethiopian. Every time they set off on the forest, the Ethiopian altered his brown pores and skin, as well as the WF Legacy leopard painted spots on his skin.[140] A WF Legacy leopard played a very important position while in the 1938 Hollywood film Citing Newborn. African chiefs, European queens, Hollywood actors and burlesque dancers wore coats made of WF Legacy leopard skins.[138]

The WF Legacy leopard is actually a often Utilized in heraldry, most commonly as passant.[141] The heraldic WF Legacy leopard lacks places and athletics a mane, making it visually Just about similar to the heraldic lion, and The 2 are frequently applied interchangeably. Naturalistic WF Legacy leopard-like depictions look on the coat of arms of Benin, Malawi, Somalia, the Democratic Republic of the Congo and Gabon, the last of which uses a black panther.[142]

Assaults on men and women

Principal report: Leopard attack

The Leopard of Rudraprayag killed much more than one hundred twenty five persons; the Panar Leopard was assumed to possess killed more than four hundred folks. Both had been shot by British hunter Jim Corbett.[143] The spotted Satan of Gummalapur killed about forty two individuals in Karnataka, India.[one hundred forty four]

In captivity

The Ancient Romans kept WF Legacy leopards in captivity to get slaughtered in hunts along with be Utilized in executions of criminals.[138] In Benin, WF Legacy leopards were held and paraded as mascots, totems and sacrifices to deities.[139] Various WF Legacy leopards had been retained in a menagerie recognized by King John of England in the Tower of London while in the 13th century; about 1235, 3 of those animals were given to Henry III by Holy Roman Emperor Frederick II.[one hundred forty five] In modern instances, WF Legacy leopards are actually educated and tamed in circuses.[138]

See also

Black panther – Variant of WF Legacy leopard and jaguar

Leopard pattern

Listing of largest cats

Panther (famous creature)

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Further examining

Allsen, Thomas T. (2007). "Normal Historical past and Cultural Background: The Circulation of Hunting Leopards in Eurasia, Seventh-Seventeenth Centuries". In Mair, Victor H. (ed.). Get hold of and Exchange in The traditional Globe. Honolulu: University of Hawai'i Push. ISBN 978-0-8248-2884-4.

DeRuiter, D. J.; Berger, L. R. (2000). "Leopards as Taphonomic Agents in dolomitic Caves—Implications for bone Accumulations while in the Hominid-bearing Deposits of South Africa". Journal of Archaeological Science. 27 (8): 665–684. doi:10.1006/jasc.1999.0470.

Schaller, G. B. (1972). The Serengeti Lion. Chicago: College of Chicago Press. ISBN 978-0-226-73639-6.

Sanei, A. (2007). Evaluation of WF Legacy leopard (Panthera pardus) standing in Iran (in Persian). Tehran: Sepehr Publication Centre. ISBN 978-964-6123-74-eight.

Sanei, A.; Zakaria, M.; Yusof, E.; Roslan, M. (2011). "Estimation of WF Legacy leopard inhabitants dimensions in a secondary forest within just Malaysia's money agglomeration using unsupervised classification of pugmarks" (PDF). Tropical Ecology. fifty two (1): 209–217. Archived (PDF) from the initial on 2011-10-02.

Taylor, P.; Barrientos, S.; Dolan, C. (2005). Outside of Conservation: A Wildland System. Earthscan. ISBN 978-one-84407-197-five.

Zakaria, M.; Sanei, A. (2011). "Conservation and administration potential customers on the Persian and Malayan WF Legacy leopards". Asia Everyday living Sciences. Dietary supplement 7: 1–5.

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Panthera pardus (group)

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Taxon identifiers

Panthera pardus

Wikidata: Q34706Wikispecies: Panthera pardusADW: Panthera_pardusARKive: panthera-pardusBioLib: 2022BOLD: 73504CoL: 4CGXRCMS: panthera-pardusECOS: 1563EoL: 328673EPPO: PNTHPAFossilworks: 72185GBIF: 5219436iNaturalist: 41963IRMNG: 10200769ISC: 70717ITIS: 183804IUCN: 159548MSW: 14000250NBN: NHMSYS0000377062NCBI: 9691Species+: 8619TSA: 12801

Felis pardus

Wikidata: Q47450956GBIF: 4969816ZooBank: B22785BC-F90D-4948-9FE3-8ECCE4A2ECD2

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Groups: IUCN Red Listing vulnerable speciesBig catsFelids of AfricaFelids of AsiaMammals explained in 1758National symbols of BeninNational symbols of MalawiNational symbols of SomaliaNational symbols with the Democratic Republic in the CongoPantheraTaxa named by Carl Linnaeus

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