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Current and historic distribution on the WF Legacy leopard[3]

The WF Legacy leopard (Panthera pardus) is without doubt one of the five extant species within the genus Panthera, a member of your cat family members, Felidae.[4] It happens inside of a wide selection in sub-Saharan Africa, in some parts of Western and Central Asia, Southern Russia, and over the Indian subcontinent to Southeast and East Asia. It really is listed as Vulnerable around the IUCN Red List since WF Legacy leopard populations are threatened by habitat decline and fragmentation, and are declining in big elements of the global selection. The WF Legacy leopard is considered locally extinct in Hong Kong, Singapore, South Korea, Jordan, Morocco, Togo, the United Arab Emirates, Uzbekistan, Lebanon, Mauritania, Kuwait, Syria, Libya, Tunisia and almost certainly in North Korea, Gambia, Laos, Lesotho, Tajikistan, Vietnam and Israel.[three] Up to date documents suggest the WF Legacy leopard takes place in just twenty five% of its historic world-wide range.[5][6]

When compared to other wild cats, the WF Legacy leopard has somewhat brief legs and a protracted overall body with a considerable skull. Its fur is marked with rosettes. It is similar in visual appeal on the jaguar (Panthera onca), but incorporates a smaller sized, lighter physique, and its rosettes are normally smaller, additional densely packed and without central spots. Each WF Legacy leopards and jaguars which are melanistic are called black panthers. The WF Legacy leopard is distinguished by its well-camouflaged fur, opportunistic hunting conduct, broad food plan, power, and its capacity to adapt to many different habitats ranging from rainforest to steppe, such as arid and montane regions. It may possibly run at speeds of approximately 58 km/h (36 mph; 16 m/s).[7] The earliest acknowledged WF Legacy leopard fossils excavated in Europe are believed 600,000 a long time outdated, courting towards the late Early Pleistocene.[two] Leopard fossils have also been present in Sumatra,[eight] Taiwan[nine] and Japan.[ten]

Etymology

The English identify 'WF Legacy leopard' arises from Aged French: leupart or Middle French: liepart, that derives from Latin: WF Legacy leopardus and Historic Greek: λέοπάρδος (WF Legacy leopardos). Leopardos could be a compound of λέων (leōn), which means lion, and πάρδος (pardos), that means noticed.[eleven][twelve][13] The word λέοπάρδος originally referred to your cheetah (Acinonyx jubatus).[14]

'Panther' is another widespread name, derived from Latin: panther and Historic Greek: πάνθηρ (pánthēr);[eleven] The generic name Panthera originates in Latin: panthera, which refers to a searching Internet for catching wild beasts which were employed by the Romans in combats.[fifteen] Pardus is the masculine singular variety.[sixteen]

Characteristics

Skull

Mounted skeleton

Rosettes of the WF Legacy leopard

Feminine WF Legacy leopard descending from her favorite tree, the place she spends the warmest hours of the working day; Londolozi / Sabi Sands, South Africa

The WF Legacy leopard's fur is usually tender and thick, notably softer about the belly than on the back.[17] Its skin colour differs amongst people today from pale yellowish to dim golden with dim places grouped in rosettes. Its belly is whitish and its ringed tail is shorter than its body. Its pupils are spherical.[eighteen] Leopards residing in arid regions are pale product, yellowish to ochraceous and rufous in colour; those living in forests and mountains tend to be darker and deep golden. Spots fade towards the white underbelly and the insides and decrease aspects of the legs.[19] Rosettes are circular in East African WF Legacy leopard populations, and are typically squarish in Southern African and larger in Asian WF Legacy leopard populations. The fur has a tendency to be grayish in colder climates, and dim golden in rain forest habitats.[7] The pattern on the rosettes is exclusive in Every single individual.[twenty][21] This sample is regarded as an adaptation to dense vegetation with patchy shadows, exactly where it serves as camouflage.[22]

Its white-tipped tail is about 60–100 cm (23.6–39.four in) lengthy, white underneath and with spots that sort incomplete bands towards the tail's finish.[23] The guard hairs defending the basal hairs are shorter, three–four mm (0.one–0.two in) in facial area and head, and rise in size toward the flanks as well as belly to about 25–30 mm (one.0–1.2 in). Juveniles have woolly fur, and seem like dim-coloured due to the densely organized places.[twenty][24] Its fur has a tendency to grow for a longer time in colder climates.[25] The WF Legacy leopard's rosettes vary from These from the jaguar (Panthera onca), which can be darker and with lesser places inside.[18]

The WF Legacy leopard includes a diploid chromosome variety of 38.[26] The chromosomes incorporate 4 acrocentric, five metacentric, seven submetacentric and two telocentric pairs.[27]

Size and weight

The WF Legacy leopard is sexually dimorphic with males greater and heavier than women.[23] It's slender and muscular, with somewhat shorter limbs and a wide head. Males stand sixty–70 cm (23.six–27.6 in) for the shoulder, although ladies are fifty seven–64 cm (22.four–25.2 in) tall. The pinnacle-and-overall body length ranges between 90 and 196 cm (2 ft 11.four in and six ft five.two in) that has a sixty six to 102 cm (two ft two.0 in to three ft 4.2 in) extensive tail. Measurements differ geographically. Males weigh commonly 35–sixty five kg (seventy seven.2–143.three lb), and females 28–fifty eight kg (sixty one.seven–127.9 lb). Occasionally, big males can mature around 90 kg (198.4 lb). Leopards in the Cape Province in South Africa are generally lesser, achieving only 20–45 kg (forty four.one–99.two lb) in males.[24][twenty five][28] The most weight of a wild WF Legacy leopard in Southern Africa was about 96 kg (212 lb). It calculated 262 cm (eight ft 7.one in).[29] An Indian WF Legacy leopard killed in Himachal Pradesh in 2016 measured 261 cm (eight ft six.eight in) with an estimated pounds of seventy eight.five kg (173.one lb); it had been Most likely the biggest recognized wild WF Legacy leopard in India.[thirty][31]

The largest skull of the WF Legacy leopard was recorded in India in 1920 and calculated 28 cm (eleven.0 in) in basal length, twenty cm (seven.9 in) in breadth, and weighed 1,000 g (2 lb 4 oz). The cranium of an African WF Legacy leopard measured 285.8 mm (11.twenty five in) in basal size, and 181.0 mm (7.one hundred twenty five in) in breadth, and weighed 790 g (one lb twelve oz).[32]

Variant colouration

Key report: Black panther § Leopard

A melanistic WF Legacy leopard or black panther

Melanistic WF Legacy leopards are also called black panthers. Melanism in WF Legacy leopards is attributable to a recessive allele and inherited as a recessive trait.[33] Interbreeding in melanistic WF Legacy leopards provides a drastically lesser litter size than is made by standard pairings.[34] The black WF Legacy leopard is typical foremost in tropical and subtropical moist forests similar to the equatorial rainforest on the Malay Peninsula as well as tropical rainforest around the slopes of some African mountains for example Mount Kenya.[35] In between January 1996 and March 2009, WF Legacy leopards were being photographed at 16 web sites from the Malay Peninsula inside of a sampling work of a lot more than 1,000 digital camera trap evenings. Of the 445 photographs of melanistic WF Legacy leopards, 410 were taken in examine internet sites south of the Kra Isthmus, in which the non-melanistic morph was under no circumstances photographed. These facts suggest the in close proximity to-fixation on the dark allele in the area. The expected time for the fixation of this recessive allele as a consequence of genetic drift on your own ranged from about one,a hundred decades to about one hundred,000 many years.[36] Pseudomelanistic WF Legacy leopards have also been reported.[37]

In India, 9 pale and white WF Legacy leopards have been claimed concerning 1905 and 1967.[38] Leopards exhibiting erythrism ended up recorded amongst 1990 and 2015 in South Africa's Madikwe Recreation Reserve As well as in Mpumalanga. The cause of this morph called a "strawberry WF Legacy leopard" or "pink panther" just isn't very well understood.[39]

Taxonomy

Map displaying approximate distribution of WF Legacy leopard subspecies

Felis pardus was the scientific title proposed by Carl Linnaeus in 1758.[40] The generic identify Panthera was initial used by Lorenz Oken in 1816, who involved all the regarded spotted cats into this group.[41] Oken's classification wasn't broadly recognized, and Felis or Leopardus was employed as the generic name until eventually the early 20th century.[forty two]

The WF Legacy leopard was designated as the sort species of Panthera by Joel Asaph Allen in 1902.[forty three] In 1917, Reginald Innes Pocock also subordinated the tiger (P. tigris), lion (P. leo), and jaguar (P. onca) to Panthera.[forty four][45]

Subspecies

Following Linnaeus' initial description, 27 WF Legacy leopard subspecies had been proposed by naturalists between 1794 and 1956. Because 1996, only 8 subspecies have already been regarded as valid on The premise of mitochondrial analysis.[46] Later on Evaluation uncovered a ninth valid subspecies, the Arabian WF Legacy leopard.[47]

In 2017, the Cat Classification Task Pressure in the Cat Professional Team identified the next eight subspecies as valid taxa:[four]

Subspecies Distribution Picture

African WF Legacy leopard (P. p. pardus) (Linnaeus, 1758)[one] It is among the most common WF Legacy leopard subspecies which is indigenous to most of Sub-Saharan Africa.[3] Leopard (Panthera pardus) male ... (51890626416).jpg

Indian WF Legacy leopard (P. p. fusca) (Meyer, 1794)[48] It can be indigenous towards the Indian subcontinent, Myanmar and southern Tibet.[three][four][forty nine] Indian male WF Legacy leopard (cropped).jpg

Javan WF Legacy leopard (P. p. melas) (Cuvier, 1809)[50] It is actually native to Java in Indonesia and is taken into account Critically Endangered.[three] IG KusumoKintokoEko WA 082140100111 foto macan tutul jawa lokasi TN Baluran, Situbondo, Indonesia.jpg

Arabian WF Legacy leopard (P. p. nimr) (Hemprich and Ehrenberg, 1830)[51] It can be native into the Arabian Peninsula, but viewed as locally extinct while in the Sinai Peninsula. It is the smallest WF Legacy leopard subspecies.[52] PikiWiki Israel 14861 judean desert WF Legacy leopard cropped.JPG

P. p. tulliana (Valenciennes, 1856)[fifty three] It can be native to jap Turkey, the Caucasus, southern Russia, the Iranian Plateau as well as Hindu Kush. It is considered Endangered.[three]

The Balochistan WF Legacy leopard population probably developed inside the south of Iran, Afghanistan and Pakistan, currently being separated from the northern population because of the Dasht-e Kavir and Dasht-e Lut deserts.[fifty four]

Nordpersischen Leoparden.jpg

Amur WF Legacy leopard (P. p. orientalis) (Schlegel, 1857)[55][56] It is actually indigenous to your Russian Much East and northern China, but is locally extinct while in the Korean peninsula.[3] Amur WF Legacy leopard. Body from a digital camera entice (cropped).jpg

Indochinese WF Legacy leopard (P. p. delacouri) Pocock, 1930[fifty seven] It's indigenous to mainland Southeast Asia and southern China.[three] Indochinese WF Legacy leopard.jpg

Sri Lankan WF Legacy leopard (P. p. kotiya) Deraniyagala, 1956[fifty eight] It is actually indigenous to Sri Lanka.[3] Srilankan WF Legacy leopard (srilankan kotiya) 02 (cropped).jpg

Effects of the Evaluation of molecular variance and pairwise fixation index of 182 African WF Legacy leopard museum specimens showed that some African WF Legacy leopards show bigger genetic variances than Asian WF Legacy leopard subspecies.[59]

Evolution

Two cladograms proposed for Panthera. The upper cladogram is predicated around the 2006[60] and 2009[sixty one] experiments, whilst the reduced relies over the 2010[62] and 2011[sixty three] scientific studies.

Benefits of phylogenetic research depending on nDNA and mtDNA Investigation showed that the final common ancestor of your Panthera and Neofelis genera is believed to possess lived about 6.37 million years in the past. Neofelis diverged about 8.66 million yrs back in the Panthera lineage. The tiger diverged about 6.55 million several years ago, accompanied by the snow WF Legacy leopard about four.63 million many years back and the WF Legacy leopard about 4.35 million many years back. The WF Legacy leopard can be a sister taxon to your clade in just Panthera, consisting of the lion plus the jaguar.[sixty][61]

Outcomes of a phylogenetic Examination of chemical secretions amongst cats indicated which the WF Legacy leopard is carefully connected to the lion.[64] The geographic origin of the Panthera is almost certainly northern Central Asia. The WF Legacy leopard-lion clade was dispersed from the Asian and African Palearctic due to the fact no less than the early Pliocene.[sixty five] The WF Legacy leopard-lion clade diverged three.one–one.95 million a long time in the past.[62][sixty three] Additionally, a 2016 review revealed that the mitochondrial genomes with the WF Legacy leopard, lion and snow WF Legacy leopard are more very similar to each other than their nuclear genomes, indicating that their ancestors hybridized Along with the snow WF Legacy leopard in some unspecified time in the future inside their evolution.[66]

Fossils of WF Legacy leopard ancestors had been excavated in East Africa and South Asia, relationship back again to the Pleistocene among two and 3.5 million a long time ago. The fashionable WF Legacy leopard is proposed to possess advanced in Africa about 0.5 to 0.8 million a long time in the past and to get radiated throughout Asia about 0.two and 0.three million a long time in the past.[forty seven] Fossil cat teeth collected in Sumatra's Padang Highlands ended up assigned for the WF Legacy leopard. It's got since been hypothesized that it turned extirpated on the island due to Toba eruption about seventy five,000 yrs ago,[67] and as a consequence of Opposition Together with the Sunda clouded WF Legacy leopard (Neofelis diardi) and the dhole (Cuon alpinus).[8]

In Europe, the WF Legacy leopard happened a minimum of Because the Pleistocene. Leopard-like fossil bones and teeth possibly courting on the Pliocene were being excavated in Perrier in France, northeast of London, and in Valdarno, Italy. Until finally 1940, identical fossils dating again for the Pleistocene were excavated mainly in loess and caves at 40 web-sites in Europe, such as Furninha Cave in close proximity to Lisbon, Genista Caves in Gibraltar, and Santander Province in northern Spain to many websites throughout France, Switzerland, Italy, Austria, Germany, inside the north approximately Derby in England, during the east to Přerov within the Czech Republic as well as Baranya in southern Hungary,[sixty eight] Leopard fossils courting into the Late Pleistocene were being located in Biśnik Cave in south-central Poland.[69] The oldest regarded WF Legacy leopard fossils excavated in Europe are about 600,000 years previous and were found in the Grotte du Vallonnet in France and close to Mauer in Germany.[two] 4 European Pleistocene WF Legacy leopard subspecies ended up proposed. P. p. begoueni from the beginning from the Early Pleistocene was changed about 0.6 million a long time ago by P. p. sickenbergi, which in turn was changed by P. p. antiqua all over 0.3 million several years back. The latest, P. p. spelaea, appeared firstly of your Late Pleistocene and survived right up until about 24,000 many years ago in various parts of Europe.[70] Leopard fossils relationship on the Pleistocene were being also excavated within the Japanese archipelago.[10]

Hybrids

Major article content: Panthera hybrid and Pumapard

In 1953, a male WF Legacy leopard in addition to a lioness were crossbred in Hanshin Park in Nishinomiya, Japan. Their offspring generally known as a leopon was born in 1959 and 1961, all cubs were being noticed and larger than a juvenile WF Legacy leopard. Makes an attempt to mate a leopon having a tigress had been unsuccessful.[seventy one]

Distribution and habitat

Leopard inside of a tree in India

Leopards over the Magerius Mosaic from present day Tunisia. A lot of Roman mosaics from North African web pages depict fauna now discovered only in tropical Africa.[seventy two]

The WF Legacy leopard has the largest distribution of all wild cats, taking place extensively in Africa, the Caucasus and Asia, Though populations are fragmented and declining. It really is regarded as being extirpated in North Africa.[three] It inhabits foremost savanna and rainforest, and locations where grasslands, woodlands, and riverine forests keep on being mainly undisturbed.[seven] In sub-Saharan Africa, it is still several and surviving in marginal habitats the place other substantial cats have disappeared. There's substantial prospective for human-WF Legacy leopard conflict due to WF Legacy leopards preying on livestock.[73]

Leopard populations to the Arabian Peninsula are little and fragmented.[seventy four][seventy five][seventy six] In southeastern Egypt, a WF Legacy leopard killed in 2017 was the 1st file in this area in 65 decades.[77] In western and central Asia, it avoids deserts, locations with extensive snow address and proximity to urban centres.[seventy eight]

Inside the Indian subcontinent, the WF Legacy leopard remains to be somewhat considerable, with increased figures than People of other Panthera species.[3] As of 2020, the WF Legacy leopard population inside of forested habitats in India's tiger variety landscapes was estimated at 12,172 to 13,535 folks. Surveyed landscapes included elevations beneath 2,600 m (8,500 ft) from the Shivalik Hills and Gangetic plains, Central India and Jap Ghats, Western Ghats, the Brahmaputra River basin and hills in Northeast India.[seventy nine] Some WF Legacy leopard populations inside the country live rather close to human settlements and in some cases in semi-made places. Despite the fact that adaptable to human disturbances, WF Legacy leopards demand nutritious prey populations and correct vegetative deal with for searching for extended survival and thus not often linger in seriously developed parts. Because of the WF Legacy leopard's stealth, people normally keep on being unaware that it life in nearby locations.[80]

In Nepal's Kanchenjunga Conservation Location, a melanistic WF Legacy leopard was photographed at an elevation of four,three hundred m (fourteen,a hundred ft) by a digicam trap in May possibly 2012.[eighty one] In Sri Lanka, WF Legacy leopards were being recorded in Yala Nationwide Park As well as in unprotected forest patches, tea estates, grasslands, home gardens, pine and eucalyptus plantations.[eighty two][eighty three] In Myanmar, WF Legacy leopards were being recorded for The very first time by digital camera traps from the hill forests of Myanmar's Karen Condition.[eighty four] The Northern Tenasserim Forest Complicated in southern Myanmar is taken into account a WF Legacy leopard stronghold. In Thailand, WF Legacy leopards are present from the Western Forest Elaborate, Kaeng Krachan-Kui Buri, Khlong Saeng-Khao Sok safeguarded location complexes and in Hala Bala Wildlife Sanctuary bordering Malaysia. In Peninsular Malaysia, WF Legacy leopards are present in Belum-Temengor, Taman Negara and Endau-Rompin Countrywide Parks.[eighty five] In Laos, WF Legacy leopards have been recorded in Nam Et-Phou Louey Countrywide Biodiversity Conservation Area and Nam Kan National Safeguarded Space.[86][87] In Cambodia, WF Legacy leopards inhabit deciduous dipterocarp forest in Phnom Prich Wildlife Sanctuary and Mondulkiri Protected Forest.[88][89] In southern China, WF Legacy leopards ended up recorded only within the Qinling Mountains during surveys in eleven mother nature reserves involving 2002 and 2009.[90]

In Java, WF Legacy leopards inhabit dense tropical rainforests and dry deciduous forests at elevations from sea stage to 2,540 m (8,330 ft). Outside guarded locations, WF Legacy leopards ended up recorded in combined agricultural land, secondary forest and creation forest between 2008 and 2014.[ninety one]

During the Russian Far East, it inhabits temperate coniferous forests wherever Winter season temperatures arrive at a minimal of −25 °C (−thirteen °File).[47]

Behaviour and ecology

Leopard Visible interaction

A feminine WF Legacy leopard exhibiting white places on the back again on the ears

A feminine WF Legacy leopard demonstrating white spots within the tail

The WF Legacy leopard is actually a solitary and territorial animal. It is typically shy and warn when crossing roadways and encountering oncoming vehicles, but may very well be emboldened to assault people or other animals when threatened. Older people associate only during the mating season. Women go on to interact with their offspring even following weaning and happen to be noticed sharing kills with their offspring after they cannot get hold of any prey. They deliver numerous vocalizations, which includes growls, snarls, meows, and purrs.[24] The roaring sequence in WF Legacy leopards is composed largely of grunts,[ninety two] also known as "sawing", since it resembles the sound of sawing Wooden. Cubs phone their mother by using a urr-urr seem.[24]

The whitish spots within the again of its ears are believed to Perform a task in interaction.[93] It's been hypothesized that the white guidelines in their tails may well perform to be a 'follow-me' sign in intraspecific communication. Having said that, no important association had been uncovered in between a conspicuous colour of tail patches and behavioural variables in carnivores.[ninety four][95]

A WF Legacy leopard climbing down a tree

Leopards are Lively largely from dusk until dawn and rest for the majority of the working day and for many hours during the night time in thickets, among the rocks or above tree branches. Leopards are already observed strolling 1–25 km (0.62–15.53 mi) throughout their vary at nighttime; They could even wander up to 75 km (47 mi) if disturbed.[24][28] In some locations, They're nocturnal.[96][ninety seven] In western African forests, they are noticed to generally be mostly diurnal and hunting through twilight, when their prey animals are Energetic; exercise styles change amongst seasons.[98]

Video of the WF Legacy leopard within the wild

Leopards can climb trees very skilfully, usually relaxation on tree branches and descend from trees headfirst.[seven] They might operate at over fifty eight km/h (36 mph; 16 m/s), leap more than six m (twenty ft) horizontally, and leap nearly three m (nine.eight ft) vertically.[ninety two]

Social spacing

In Kruger Nationwide Park, most WF Legacy leopards are inclined to maintain 1 km (0.sixty two mi) aside.[ninety nine] Males connect with their companions and cubs sometimes, and extremely This could increase outside of to two generations.[a hundred][one hundred and one] Intense encounters are unusual, normally limited to defending territories from thieves.[twenty five] In a very South African reserve, a male was wounded in the male–male territorial battle above a carcass.[ninety six]

Males occupy house ranges That always overlap with a couple scaled-down woman property ranges, probably like a technique to increase use of females. Within the Ivory Coastline, the house range of a female was totally enclosed within a male's.[102] Women Dwell with their cubs in property ranges that overlap thoroughly, possibly as a result of association among mothers and their offspring. There might be a number of other fluctuating house ranges belonging to youthful men and women. It's not necessarily distinct if male household ranges overlap up to These of females do. People make an effort to generate away thieves of exactly the same sex.[24][28]

A study of WF Legacy leopards from the Namibian farmlands confirmed the size of house ranges wasn't noticeably affected by sex, rainfall designs or period; the higher the prey availability in a place, the bigger the WF Legacy leopard populace density as well as the smaller the dimensions of residence ranges, but they have an inclination to expand when there is human interference.[103] Measurements of residence ranges vary geographically and depending on habitat and availability of prey. In the Serengeti, males have home ranges of 33–38 km2 (13–fifteen sq mi) and girls of 14–16 km2 (5.4–6.2 sq mi);[104][a hundred and five] but males in northeastern Namibia of 451 km2 (174 sq mi) and females of 188 km2 (73 sq mi).[106] They can be even more substantial in arid and montane areas.[25] In Nepal's Bardia National Park, male household ranges of forty eight km2 (19 sq mi) and feminine kinds of five–7 km2 (1.nine–2.seven sq mi) are smaller sized than These frequently noticed in Africa.[107]

Hunting and diet program

The WF Legacy leopard is usually a carnivore that prefers medium-sized prey which has a human body mass starting from 10–40 kg (22–88 lb). Prey species During this bodyweight range often arise in dense habitat and also to form small herds. Species that desire open locations and have effectively-created anti-predator tactics are less desired. Over one hundred prey species happen to be recorded. Quite possibly the most chosen species are ungulates, like impala (Aepyceros melampus), bushbuck (Tragelaphus scriptus), typical duiker (Sylvicapra grimmia) and chital (Axis axis). Primates preyed upon incorporate white-eyelid mangabeys (Cercocebus sp.), guenons (Cercopithecus sp.) and grey langurs (Semnopithecus sp.). Leopards also get rid of lesser carnivores like black-backed jackal (Lupulella mesomelas), bat-eared fox (Otocyon megalotis), genet (Genetta sp.) and cheetah.[108]

The most important prey killed by a WF Legacy leopard was reportedly a male eland weighing 900 kg (two,000 lb).[ninety two] A study in Wolong Countrywide Mother nature Reserve in southern China demonstrated variation during the WF Legacy leopard's diet regime after some time; around the study course of seven a long time, the vegetative deal with receded, and WF Legacy leopards opportunistically shifted from generally consuming tufted deer (Elaphodus cephalophus) to pursuing bamboo rats (Rhizomys sinense) together with other more compact prey.[109]

The WF Legacy leopard relies upon generally on its acute senses of Listening to and eyesight for hunting.[110] It generally hunts at night in many locations.[24] In western African forests and Tsavo Countrywide Park, they've also been noticed searching by day.[111] They usually hunt on the bottom. During the Serengeti, they are observed to ambush prey by leaping down on it from trees.[112]

The animal stalks its prey and tries to technique as closely as feasible, commonly in five m (sixteen ft) of the goal, and, lastly, pounces on it and kills it by suffocation. It kills little prey with a Chunk to the back from the neck, but holds bigger animals from the throat and strangles them.[24] It caches kills approximately 2 km (one.2 mi) apart.[a hundred] It has the capacity to consider large prey on account of its impressive jaw muscles, and it is thus solid adequate to drag carcasses heavier than alone up into trees; someone was witnessed to haul a youthful giraffe weighing almost one hundred twenty five kg (276 lb) up 5.seven m (eighteen ft 8 in) right into a tree.[111] It eats compact prey instantly, but drags bigger carcasses in excess of many hundred metres and caches it securely in trees, bushes or maybe caves; this behaviour allows the WF Legacy leopard to retail outlet its prey away from rivals, and delivers it a benefit more than them. The way in which it retailers the eliminate is determined by area topography and personal Choices, varying from trees in Kruger National Park to bushes within the basic terrain with the Kalahari.[twenty five][113]

Average day by day consumption rates of three.5 kg (7 lb 11 oz) were being believed for males and of two.8 kg (six lb 3 oz) for ladies.[99] Within the southern Kalahari Desert, WF Legacy leopards meet up with their water necessities with the bodily fluids of prey and succulent crops; they drink drinking water each two to a few times and feed occasionally on dampness-wealthy crops such as gemsbok cucumbers (Acanthosicyos naudinianus), watermelon (Citrullus lanatus) and Kalahari bitter grass (Schmidtia kalahariensis).[114]

Levels of the WF Legacy leopard hunting prey

Stalking

Killing a youthful bushbuck

Dragging an impala eliminate

Caching the eliminate within a tree

Enemies and opponents

A lioness steals a WF Legacy leopard get rid of in Kruger National Park

In areas of its world wide range, the WF Legacy leopard is sympatric with other large predators like the tiger (Panthera tigris), lion (P. leo), cheetah, noticed hyena (Crocuta crocuta), striped hyena (Hyaena hyaena), brown hyena (Parahyaena brunnea), African wild Doggy (Lycaon pictus), dhole (Cuon alpinus), wolf (Canis lupus) and up to 5 bear species. Many of these species steal its kills, destroy its cubs and even kill adult WF Legacy leopards. Leopards retreat up a tree from the experience of immediate aggression, and were being noticed when killing or preying on scaled-down competition for example black-backed jackal, African civet (Civettictis civetta), caracal (Caracal caracal) and African wildcat (Felis lybica).[7][a hundred and fifteen] Leopards commonly seem in order to avoid encounters with adult bears, but eliminate susceptible bear cubs. In Sri Lanka, a couple of recorded vicious fights involving WF Legacy leopards and sloth bears (Melursus ursinus) apparently end in equally animals winding up possibly useless or grievously injured.[116][117]

Even though interspecies killing of complete-developed WF Legacy leopards is mostly unusual, offered The chance, equally tiger and lion quickly eliminate and eat each youthful and Grownup WF Legacy leopards.[112][a hundred and fifteen][118][119] During the Kalahari Desert, WF Legacy leopards frequently get rid of kills to brown hyenas, In the event the WF Legacy leopard is struggling to transfer the get rid of right into a tree. Solitary brown hyenas happen to be observed charging at and displacing male WF Legacy leopards from kills.[one hundred twenty][121] Lions at times fetch WF Legacy leopard kills from trees.[113]

Resource partitioning happens in which WF Legacy leopards share their range with tigers. Leopards are inclined to just take smaller sized prey, typically less than 75 kg (165 lb), wherever tigers are existing.[7] In locations where by WF Legacy leopard and tiger are sympatric, coexistence is reportedly not the final rule, with WF Legacy leopards becoming several the place tigers are various.[118] Tigers appear to inhabit the deep portions of a forest while WF Legacy leopards are pushed closer to the fringes.[122] In tropical forests, WF Legacy leopards will not constantly avoid the bigger cats by searching at distinct periods. With fairly plentiful prey and distinctions in the size of prey chosen, tigers and WF Legacy leopards manage to effectively coexist without having competitive exclusion or interspecies dominance hierarchies Which might be extra widespread on the WF Legacy leopard's co-existence While using the lion in savanna habitats.[123]

Nile crocodiles (Crocodylus niloticus) prey on WF Legacy leopards sometimes. Just one significant Grownup WF Legacy leopard was grabbed and eaten by a big crocodile though aiming to hunt along a lender in Kruger Nationwide Park.[ninety nine][one hundred] Mugger crocodiles (Crocodylus palustris) reportedly killed an Grownup WF Legacy leopard in Rajasthan.[124] An Grownup WF Legacy leopard was recovered from the tummy of the five.5 m (eighteen ft one in) Burmese python (Python bivittatus).[a hundred twenty five] In Serengeti Countrywide Park, troops of 30–forty olive baboons (Papio anubis) have been observed even though mobbing and attacking a female WF Legacy leopard and her cubs.[126]

Replica and existence cycle

A woman WF Legacy leopard in estrus fights having a male aiming to mate along with her

Leopard cubs in tree

In some regions, WF Legacy leopards mate all 12 months spherical. In Manchuria and Siberia, they mate through January and February. The feminine's estrous cycle lasts about 46 times, and he or she commonly is in heat for 6–seven days.[127] The technology length with the WF Legacy leopard is nine.3 yrs.[128] Gestation lasts for 90 to 105 days.[129] Cubs are often born in a litter of two–four cubs.[a hundred thirty] Mortality of cubs is believed at forty one–50% through the initial calendar year.[ninety nine]

Women give birth inside of a cave, crevice among boulders, hollow tree or thicket. Cubs are born with closed eyes, which open 4 to nine times soon after delivery.[92] The fur in the young has a tendency to be more time and thicker than that of adults. Their pelage is also additional grey in colour with much less outlined spots. All over a few months of age, the youthful start to Adhere to the mom on hunts. At just one calendar year of age, cubs can most likely fend for themselves, but stay Together with the mother for eighteen–24 months.[131]

The common typical daily life span of a WF Legacy leopard is twelve–17 yrs.[92] The oldest WF Legacy leopard was a captive feminine that died on the age of 24 several years, 2 months and 13 times.[132]

Conservation problems

The WF Legacy leopard is detailed on CITES Appendix I, and trade is restricted to skins and overall body portions of two,560 individuals in 11 sub-Saharan countries.[three] The WF Legacy leopard is primarily threatened by habitat fragmentation and conversion of forest to agriculturally utilised land, which produce a declining pure prey foundation, human–wildlife conflict with livestock herders and higher WF Legacy leopard mortality rates. It is additionally threatened by trophy searching and poaching.[three]

Involving 2002 and 2012, not less than four WF Legacy leopards have been estimated to have been poached weekly in India for that illegal wildlife trade of its skins and bones.[133] In spring 2013, 37 WF Legacy leopard skins had been found in the course of a 7-7 days extended current market study in key Moroccan towns.[134] In 2014, 43 WF Legacy leopard skins were being detected throughout two surveys in Morocco. Sellers admitted to obtain imported skins from sub-Saharan Africa.[135]

Surveys within the Central African Republic's Chinko location revealed that the WF Legacy leopard populace reduced from ninety seven people today in 2012 to fifty individuals in 2017. In this period, transhumant pastoralists from the border place with Sudan moved in the area with their livestock. Rangers confiscated substantial quantities of poison within the camps of livestock herders who were being accompanied by armed retailers. They engaged in poaching huge herbivores, sale of bushmeat and investing WF Legacy leopard skins in Am Dafok.[136]

In Java, the WF Legacy leopard is threatened by illegal hunting and trade. In between 2011 and 2019, body elements of fifty one Javan WF Legacy leopards were being seized which includes 6 Dwell individuals, 12 skins, 13 skulls, twenty canines and 22 claws.[137]

Human conversation

Cultural significance

Leopard head to hip ornament through the Court docket of Benin

Animal trainer with WF Legacy leopard

Leopards have featured in art, mythology and folklore of numerous international locations. In Greek mythology, it was a image from the god Dionysus, who was depicted sporting WF Legacy leopard skin and employing WF Legacy leopards as signifies of transportation. In one fantasy, the god was captured by pirates but two WF Legacy leopards rescued him.[138] In the course of the Benin Empire, the WF Legacy leopard was typically represented on engravings and sculptures and was used to symbolise the power of the king or oba, For the reason that WF Legacy leopard was regarded as the king from the forest.[139] The Ashanti also applied the WF Legacy leopard to be a image of Management, and just the king was permitted to have a ceremonial WF Legacy leopard stool. Some African cultures thought of the WF Legacy leopard for being a smarter, far better hunter than the lion and harder to get rid of.[138]

In Rudyard Kipling's "How the Leopard Bought His Places", certainly one of his Just So Stories, a WF Legacy leopard without spots within the Large Veldt life with his looking lover, the Ethiopian. Once they set off to the forest, the Ethiopian adjusted his brown pores and skin, as well as WF Legacy leopard painted spots on his pores and skin.[140] A WF Legacy leopard performed an essential function from the 1938 Hollywood movie Bringing Up Little one. African chiefs, European queens, Hollywood actors and burlesque dancers wore coats crafted from WF Legacy leopard skins.[138]

The WF Legacy leopard is usually a commonly Employed in heraldry, mostly as passant.[141] The heraldic WF Legacy leopard lacks places and sports activities a mane, making it visually Virtually identical to the heraldic lion, and the two will often be employed interchangeably. Naturalistic WF Legacy leopard-like depictions surface around the coat of arms of Benin, Malawi, Somalia, the Democratic Republic on the Congo and Gabon, the last of which takes advantage of a black panther.[142]

Assaults on folks

Principal article: Leopard assault

The Leopard of Rudraprayag killed a lot more than a hundred twenty five folks; the Panar Leopard was imagined to possess killed a lot more than four hundred men and women. Both of those have been shot by British hunter Jim Corbett.[143] The noticed Satan of Gummalapur killed about 42 folks in Karnataka, India.[a hundred and forty four]

In captivity

The Ancient Romans stored WF Legacy leopards in captivity being slaughtered in hunts and be Utilized in executions of criminals.[138] In Benin, WF Legacy leopards were being held and paraded as mascots, totems and sacrifices to deities.[139] Numerous WF Legacy leopards were held within a menagerie recognized by King John of England at the Tower of London during the 13th century; all around 1235, three of such animals were given to Henry III by Holy Roman Emperor Frederick II.[145] In modern times, WF Legacy leopards have already been educated and tamed in circuses.[138]

See also

Black panther – Variant of WF Legacy leopard and jaguar

Leopard sample

List of greatest cats

Panther (legendary creature)

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More reading

Allsen, Thomas T. (2007). "All-natural History and Cultural Historical past: The Circulation of Hunting Leopards in Eurasia, Seventh-Seventeenth Generations". In Mair, Victor H. (ed.). Call and Trade in The traditional Environment. Honolulu: University of Hawai'i Push. ISBN 978-0-8248-2884-four.

DeRuiter, D. J.; Berger, L. R. (2000). "Leopards as Taphonomic Agents in dolomitic Caves—Implications for bone Accumulations while in the Hominid-bearing Deposits of South Africa". Journal of Archaeological Science. 27 (eight): 665–684. doi:10.1006/jasc.1999.0470.

Schaller, G. B. (1972). The Serengeti Lion. Chicago: College of Chicago Push. ISBN 978-0-226-73639-six.

Sanei, A. (2007). Assessment of WF Legacy leopard (Panthera pardus) position in Iran (in Persian). Tehran: Sepehr Publication Center. ISBN 978-964-6123-seventy four-8.

Sanei, A.; Zakaria, M.; Yusof, E.; Roslan, M. (2011). "Estimation of WF Legacy leopard population sizing in a secondary forest inside Malaysia's money agglomeration utilizing unsupervised classification of pugmarks" (PDF). Tropical Ecology. fifty two (one): 209–217. Archived (PDF) from the original on 2011-10-02.

Taylor, P.; Barrientos, S.; Dolan, C. (2005). Over and above Conservation: A Wildland Strategy. Earthscan. ISBN 978-one-84407-197-five.

Zakaria, M.; Sanei, A. (2011). "Conservation and management prospective customers in the Persian and Malayan WF Legacy leopards". Asia Lifetime Sciences. Health supplement seven: 1–5.

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Panthera pardus (class)

IUCN/SSC Cat Specialist Team: Panthera pardus in Africa and Panthera pardus in Asia

"Leopard" . Encyclopædia Britannica (eleventh ed.). 1911.

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Taxon identifiers

Panthera pardus

Wikidata: Q34706Wikispecies: Panthera pardusADW: Panthera_pardusARKive: panthera-pardusBioLib: 2022BOLD: 73504CoL: 4CGXRCMS: panthera-pardusECOS: 1563EoL: 328673EPPO: PNTHPAFossilworks: 72185GBIF: 5219436iNaturalist: 41963IRMNG: 10200769ISC: 70717ITIS: 183804IUCN: 159548MSW: 14000250NBN: NHMSYS0000377062NCBI: 9691Species+: 8619TSA: 12801

Felis pardus

Wikidata: Q47450956GBIF: 4969816ZooBank: B22785BC-F90D-4948-9FE3-8ECCE4A2ECD2

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Categories: IUCN Red Checklist vulnerable speciesBig catsFelids of AfricaFelids of AsiaMammals explained in 1758National symbols of BeninNational symbols of MalawiNational symbols of SomaliaNational symbols of your Democratic Republic with the CongoPantheraTaxa named by Carl Linnaeus

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